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Explanation:

The first clue is that sentence 5 starts the paragraph as it introduces the theme and that it is followed by sentence 4. This gives us the beginning of the paragraph as 5-4. After this, the question becomes difficult as the link to the next sentence is not very clear. However, sentence 1 comes immediately after the 5-4 pair because “the epidemic that swept through Europe” in sentence 5 is now continued in sentence 1 after the interruption of sentence 4. Sentence 4 cannot be placed anywhere else in the paragraph. So, we get the 5-4-1 sequence. After this, it is easy to see that sentence 2 and 3 come next in that order. Sentence 2 states that the oldest sample was of 5000 years ago and sentence 3 says that this age corresponds to a time of mass exodus.
Hence, the correct answer is 54123.

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